Prof. Dr. Martin Entling



Fortstr. 7
Gebäude: I
Raum: 3.08
76829 Landau


Tel.: +49 6341 280-31537

Fax: +49 6341 280-31185



Bis auf weiteres finden keine Sprechstunden statt, Anfragen bitte per mail.

since 2010Head of Ecosystem Analysis lab at University of Koblenz-Landau
2014-2018Head of the Institute for Environmental Sciences
2010Venia Docendi in Ecology, University of Bern
2009CAS in Higher Education, University of Bern
2004-2010Research and teaching at University of Bern
2004PhD in Agricultural Sciences, University of Göttingen
2000Diploma in Biology, University of Göttingen
  • Trophic interactions
  • Landscape ecology
  • Ecosystem services
  • Biodiversity conservation
  • Spider ecology


Reiff JM, Pennington T, Kolb S, ... Entling MH, Hoffmann C (2023). Consistent benefits of fungicide reduction on arthropod predators and predation rates in viticulture: a five-year experiment. BioControl 1-15.

Reiff JM, Sudarsan K, Hoffmann C, Entling MH (2023). Arthropods on grapes benefit more from fungicide reduction than from organic farming. Pest Management Science.

Beaumelle L, Giffard B, Tolle P, Winter S, Entling MH, Benítez E, Zaller JG, Auriol A, Bonnard O, Charbonnier Y, Fabreguettes O, Joubard B, Kolb S, Ostandie N, Reiff JM, Richart-Cervera S, Rusch A (2023). Biodiversity conservation, ecosystem services and organic viticulture: A glass half-full. Agriculture, Ecosystems & Environment 351: 108474.


Narimanov N, Bonte D & Entling MH (2022) Heritability of dispersal in a rapidly spreading invasive spider in Europe. Animal Behaviour 183: 93-101.

Ammann L, Bosem-Baillod A, Eckerter PW, Entling MH, Albrecht M & Herzog F (2022) Comparing floral resource maps and land cover maps to predict predators and aphid suppression on field bean. Landscape Ecology 37: 431-441.

González E, Felix B, Eckerter PW, Pfaff V, Weiler S & Entling MH (2022) Ecological requirements drive pest and natural enemy response to landscape context in wheat. Journal of Applied Ecology 59: 444-456.

Allen-Perkins A, Magrach A, Dainese M, Garibaldi LA, Kleijn D, Rader R, Reilly JR, Winfree R, Lundin O, McGrady CM, …, Entling MH, …, Bartomeus I (2022) CropPol: a dynamic, open and global database on crop pollination. Ecology 103: e3614.

Buse J, Reiff D & Entling MH (2022) Ground beetles (Coleoptera: Carabidae) on a recently established wood-pasture: a case study from southwest Germany. Angewandte Carabidologie 14: 32-39.

Kaczmarek M, Entling MH & Hoffmann C (2022) Using Malaise Traps and Metabarcoding for Biodiversity Assessment in Vineyards: Effects of Weather and Trapping Effort. Insects13: 507.

Eckerter PW, Albrecht M, Bertrand C, Gobet E, Herzog F, Pfister SC, Tinner W & Entling MH (2022) Efects of temporal foral resource availability and non‑crop habitats on broad bean pollination. Landscape Ecology 37: 1573-1586.

Wersebeckmann V, Entling MH & Leyer I (2022) Revegetation of vineyard terrace embankments: a matter of seed mixture and seeding technique. Journal of Environmental Management 317: 115409.

Schirmel J, Petschner S & Entling MH (2022) Positive effects of organic viticulture on carabid beetles depend on landscape and local habitat conditions. Annals of Applied Biology 181: 192-200.

Laux M, Lyu H, Entling MH, Schirmel J, Narang A, Köhler M & Saha S (2022) Native pedunculate oaks support more biodiversity than non-native oaks, but non-native oaks are healthier than native oaks: A study on street and park trees of a city. Science of the Total Environment 853: 158603.

Eckerter PW, Albrecht M, Herzog F & Entling MH (2022) Flower availability in agricultural landscapes, resource use and fitness consequences for two species of wild bees. Basic and Applied Ecology 65: 1-15.

Gerstle V, Manfrin A, Kolbenschlag S, Gerken M, Mufachcher Ul Islam ASM, Entling MH, Bundschuh M & Brühl C (2022) Benthic macroinvertebrate community shifts based on Bti-induced chironomid reduction also decrease Odonata emergence. Environmental Pollution 316: 120488.

González E, Bianchi FJJA, Wizorek S, Schumacher M & Entling MH (2022) Variation in pollen digestion times by the green lacewing Chrysoperla carnea (Stephens) between plant species. Entomologia Experimentalis et Applicata 170: 1049-1054.

Narimanov N, Bauer T, Bonte D, Fahse L & Entling MH (2022) Accelerated invasion through the evolution of dispersal behavior. Global Ecology and Biogeography 31: 2423-2436.


Herbertsson L, Ekroos J, Albercht M, Bartomeus I, Batáry P, Bommarco R, Caplat P, Diekötter T, Eikestam JM, Entling MH, Farbu S, Farwig N, Gonzalez-Varo JP, Hass A, Holzschuh A, Hopfenmüller S, Jakobsson A, Jauker B, Kovács-Hostyánszki A, Kunin WE, Lindström S, Mullen S, Öckinger E, Pentanidou T, Potts SG, Power EF, Rundlöf M, Seibel K, Sober V, Sönderman A, Steffan-Dewenter I, Stout J, Teder T, Tscharntke T, Winsa M & Smith HG (2021) Bees increase seed set of wild plants while increasing proportion of arable land has a variable effect on pollination in European agricultural landscapes. Plant Ecology and Evolution 154: 341-350.

Hussain RI, Brandl M, Maas B, Rabl D, Walcher R, Krautzer B, Entling MH, Moser D & Frank T (2021) Re-established grasslands on farmland promote pollinators more than predators. Agriculture, Ecosystems and Environment 319: 107543.

Narimanov N, Bonte D, Mason P, Mestre L & Entling MH (2021) Disentangling the role of electric fields and wind in aerial dispersal. Journal of Arachnology 49: 380-383.

Narimanov N, Hatamli K & Entling MH (2021) Prey naïveté rather than enemy release dominates the relation of an invasive spider toward a native predator. Ecology and Evolution 11: 11200-11206.

Narimanov N, Kempel A, van Kleunen M & Entling MH (2021) Unexpected sensitivity of the highly invasive spider Mermessus trilobatus to soil disturbance in grasslands. Biological Invasions 23: 1-6.

Reiff JM, Ehringer M, Hoffmann C & Entling MH (2021) Fungicide reduction favors the natural control of pest mites under both organic and conventional viticulture. Agriculture, Ecosystems and Environment 305: 107172.

Reiff J, Kolb S, Entling MH, Herndl T, Möth S, Walzer A, Kropf M & Winter S (2021) Organic farming and cover-crop management reduce pest predation in Austrian vineyards. Insects 12: 220.

Rösch V, Aloisio P & Entling MH (2021) Prey, management and landscape requirements of an endangered population of the woodlark Lullula arborea in Southwest Germany. Journal of Ornithology 162:689-698.

Wersebeckmann V, Kolb S, Entling MH & Leyer I (2021) Maintaining steep slope viticulture for spider diversity. Global Ecology and Conservation 29: e01727.


Kolb S, Uzman D, Leyer I, Reineke A & Entling MH (2020) Differential effects of semi-natural habitats and organic management on spiders in viticultural landscapes. Agriculture, Ecosystems and Environment 287: 106695.

Graf N, Battes KP, Cimpean M, Entling MH, Frisch K, Link M, Scharmüller A, Schreiner VC, Szöcs E, Zubrod JP & Schäfer RB (2020) Relationship between agricultural pesticides and the diet of riparian spiders in the field. Environmental Sciences Europe 32: 1.

Buse J & Entling MH (2020) Stronger dung removal in forests compared with grassland is driven by trait composition and biomass of dung beetles. Ecological Entomology 45: 223-231.

Uzman D, Reineke A, Entling MH & Leyer I (2020) Habitat area and connectivity support cavity-nesting bees in vineyards more than organic management. Biological Conservation 242: 108419.

Picchi MS, Bocci G, Petacchi R & Entling MH (2020) Taxonomic and functional differentiation of spiders in habitats in traditional olive production landscapes in Italy. European Journal of Entomology 117: 18-26.

Mestre L, Menzel F & Entling MH (2020) Non-consumptive effects between predators depend on the foraging mode of intraguild prey. Journal of Animal Ecology 89: 1690-1700.

Holland JM, Jeanneret P, Moonen A-C, van der Werf W, Rossing WAH, Antichi D, Entling MH, Giffard B, Helsen H, Szalai M, Rega C, Gibert C & Veroman E (2020) Approaches to identify the value of seminatural habitats for conservation biological control. Insects 11: 195.

Holland JM, Sutter L, Albrecht M, Jeanneret P, Pfister SC, Schirmel J, Entling MH, Kaasik R, Kovacs G, Veromann E, Bartual AM, Marini S, Moonen A-C, Szalai M, Helsen H, Winkler K, Lof ME, van der Werf W, McHugh NM, Smith BM, Wallis DW  & Cresswell JE (2020) Moderate pollination limitation in some entomophilous crops of Europe. Agriculture, Ecosystems and Environment 302: 107002.

Ammann L, Moorhouse-Gann R, Bertrand C, Mestre L, Cuff J, Pérez Hidalgo N, Ellison A, Herzog F, Entling MH, Albrecht M & Symondson WOC (2020) Insights into aphid prey consumption by ladybirds: Optimising field sampling methods and primer design for High Throughput Sequencing. PLOS ONE 15: e0235054.

Albrecht M, Kleijn D, Williams NM, Tschumi M, Blaauw BR, Bommarco R, Campbell AJ, Dainese M, Drummond FA, Entling MH, Ganser D, De Groot GA, Goulson D, Grab H, Hamilton H, Herzog F, Isaacs R, Jacot K, Jeanneret P, Jonsson M, Knop E, Kremen C, Landis DA, Loeb GM, Marini L, McKerchar M, Morandin L, Pfister SC, Potts SG, Rundlöf M, Sardinas H, Sciligo A, Thies C, Tscharntke T, Venturini E, Veroman E, Vollhardt IMG, Wäckers F, Ward K, Wilby A, Woltz M, Wratten S & Sutter L (2020) The effectiveness of flower strips and hedgerows on pestcontrol, pollination services and crop yield: a quantitative synthesis. Ecology Letters 23: 1488-1498.

Uzman D, Entling MH, Leyer I & Reineke A (2020) Mutual and opposing responses of carabid beetles and predatory wasps to local and landscape factors in vineyards. Insects 11: 746.

Eckerter PW, Albus L, Natarajan S, Albrecht M, Ammann L, Gobet E, Herzog F, Tinner W & Entling MH (2020) Using temporally resolved floral resource maps to explain bumblebee colony performance in agricultural landscapes. Agronomy 10: 1993.


Michalko R, Pekár S & Entling MH (2019) An updated perspective on spiders as generalist predators in biological control. Oecologia 189: 21-36.

Uzman D, Pliester J, Leyer I, Entling MH & Reineke A (2019) Drivers of entomopathogenic fungi presence in organic and conventional vineyard soils. Applied Soil Ecology 133: 89-97.

Graf N, Battes KP, Cimpean M, Dittrich P, Entling MH, Link M, Scharmüller A, Schreiner VC, Szöcs E & Schaefer RB (2019) Do agricultural pesticides in streams affect riparian spiders? Science of the Total Environment 66: 126-135.

Bartual AM, Sutter L, Bocci G, Moonen A-C, Cresswell J, Entling M, Giffard B, Jacot K, Jeanneret P, Holland J, Pfister S, Pintér O, Veromann E, Winkler K & Albrecht M (2019) The potential of different semi-natural habitats to sustain pollinators and natural enemies in European agricultural landscapes. Agriculture, Ecosystems and Environment 279: 43-52.

Michalko R, Pekár S, Dul’a M & Entling MH (2019) Global patterns in the biocontrol efficacy of spiders: a meta-analysis. Global Ecology and Biogeography 28:1366–1378.

Martin EA, Dainese M, Clough Y, Báldi A, Bommarco R, Gagic V, Garratt MP, Holzschuh A, Kleijn D, Kovács‐Hostyánszki A, Marini L, … , Entling MH, … , Steffan-Dewenter I (2019) The interplay of landscape composition and configuration: new pathways to manage functional biodiversity and agro-ecosystem services across Europe. Ecology Letters 22: 1083-1094.

Pennington T, Kolb S, Kaiser J, Hoffmann C & Entling MH (2019) Do minimal pruning and reduced plant protection enhance spiders in vineyards? Journal of Arachnology 47: 381–384.

Rösch V, Hoffmann M, Diehl U & Entling MH (2019) Newly created wood pastures as a tool for bird and grasshopper conservation. Biological Conservation 237: 493-503.

Bertrand C, Eckerter P, Ammann L, Entling MH, Gobet E, Herzog F, Mestre L, Tinner W & Albrecht M (2019) Seasonal shifts and complementary use of pollen sources by key pollinators and aphid enemies in European agricultural landscapes. Journal of Applied Ecology 56: 2431–2442.


Fusser MS, Holland JM, Jeanneret P, Pfister SC, Entling MH & Schirmel J (2018) Interactive effects of local and landscape factors on farmland carabids. Agricultural and Forest Entomology 20: 549-557.

Karp DS, Chaplin-Kramer R, Meehan TD, Martin EA, DeClerck F, ..., Entling MH, ..., Zou Y (2018) Crop pests and predators exhibit inconsistent responses to surrounding landscape composition. PNAS 115: E7863-E7870.

Mestre L, Hetz J, Kolb S, Pfister SC, Amato M, Sutter L, Jeanneret P, Albrecht M, Schirmel J & Entling MH (2018) Both woody and herbaceous semi-natural habitats are essential for spider overwintering in European farmland. Agriculture, Ecosystems and Environment 267: 141-146.

Pennington T, Reiff JM, Entling MH & Hoffmann C (2018) Reduced plant protection in fungus resistant cultivars favors natural control of a major grapevine pest. Biocontrol 63: 687-695.

Pfister SC, Eckerter PW, Krebs J, Cresswell JE, Schirmel J & Entling MH (2018) Dominance of cropland reduces the pollen deposition from bumble bees. Scientific Reports 8: 13873.

Schirmel J, Albrecht M, Bauer P-M, Sutter L, Pfister SC & Entling MH (2018) Semi-natural habitats in the surrounding landscape promote hover flies in agroecosystems of two European countries. Journal of Applied Ecology 55: 1747-1758.


Fusser MS, Pfister SC, Entling MH & Schirmel J (2017) Effects of field margin type and landscape composition on predatory carabids and slugs in wheat fields. Agriculture, Ecosystems and Environment 247: 182-188.

Graf N, Bucher R, Schaefer RB & Entling MH (2017) Contrasting effects of aquatic subsidies on a terrestrial trophic cascade. Biology Letters 13: 20170129.

Hoffmann C, Köckerling J, Biancu S, Gramm T, Michl G & Entling MH (2017) Can flowering greencover crops promote biological control in German vineyards? Insects 8: 121.

Hudson L, Newbold T, Contu S, Hill SLL, Lysenko I, ..., Entling MH, ..., Ewers RM, Mace GM, Purves DW, Scharlemann JPW & Purvis A (2017) The database of the PREDICTS (Projecting Responses of Ecological Diversity in Changing Terrestrial Systems) project. Ecology and Evolution 7: 145-188.

Pennington T, Kraus C, Alakina E, Entling MH & Hoffmann C (2017) Minimal pruning and reduced plant protection promote predatory mites in grapevine. Insects 8: 86.

Pfister SC, Sutter L, Albrecht M, Marini S, Schirmel J & Entling MH (2017) Positive effects of local and landscape features on predatory flies in Europen agricultural landscapes. Agriculture, Ecosystems and Environment 239: 283-292.

Pfister SC, Eckerter PW, Schirmel J, Cresswell JE & Entling MH (2017) Sensitivity of commercial pumpkin yield to potential decline among different groups of pollinating bees. Royal Society Open Science 4: 170102.

Pfister SC, Schirmel J & Entling MH (2017) Aphids and their enemies in pumpkin respond differently to management, local and landscape features. Biological Control 115: 37-45.

Stiegel S, Entling MH & Mantilla-Contreras J (2017) Reading the leaves’ palm: leaf traits and herbivory along the microclimatic gradient of forest layers. PLOS ONE 12: e0169741.

Wendland S, Baudy P, Fusser MS, Entling MH & Schirmel J (2017) Contrasting predation pressure of different ground beetles on slugs and their eggs. Biocontrol 62: 749-756.


Binz H, Kraft EF, Entling MH & Menzel F (in press) Behavioral response of a generalist predator to chemotactile cues of two taxonomically distinct prey species. Chemoecology 26: 153-162.

Bucher R, Wedel MF, Entling MH, Andres C & Nickel H (2016) Biodiversity in low-intensity pastures, straw meadows, and fallows of a fen area – a multitrophic comparison. Agriculture, Ecosystems and Environment 219: 190-196.

Buri P, Humbert J-Y, Stanska M, Hajdamowicz I, Tran E, Entling MH & Arlettaz R (2016) Delayed mowing promotes planthoppers, leafhoopers and spiders in extensively managed meadows. Insect Conservation and Diversity 9: 536-545.

Buse J, Entling MH, Ranius T & Assmann T (2016) Response of saproxylic beetles to small-scale habitat connectivity depends on trophic levels. Landscape Ecology 31: 939-949.

Coudrain V, Rittiner S, Herzog F, Tinner W & Entling MH (2016) Landscape distribution of food and nesting sites affect larval diet and nest size, but not abundance of Osmia bicornis. Insect Science 23: 746-753.

De Palma A, Abrahamczyk S, Albrecht M, Baset Y, Blake RJ, Bugter R, Connop S, Cunnigham SA, Darvill B, Diekötter T, Downing N, Entling MH, ... & Purvis A (2016) ) Predicting bee community responses to land-use changes: Effects of geographic and taxonomic biases. Scientific Reports 6: 31153.

Fusser M, Pfister S, Entling MH & Schirmel J (2016) Effects of landscape composition on carabids and slugs in herbaceous and woody field margins. Agriculture, Ecosystems and Environment 226: 79-87.

Holland JM, Bianchi FJJA, Entling MH, Moonen A-C, Smith BM & Jeanneret P (2016) Structure, function and management of semi-natural habitats for conservation biological control: A review of European studies. Pest Management Science 72: 1638-1651.

Müller IB, Buhk C, Alt M, Entling MH & Schirmel J (2016) Plant functional shifts in Central European grassland under traditional flood irrigation. Applied Vegetation Science 19: 122-131.

Müller IB, Buhk C, Lange D, Entling MH & Schirmel J (2016) Contrasting effects of irrigation and fertilization on plant diversity in hay meadows. Basic and Applied Ecology 17: 576-585.

Picchi M, Bocci G, Petacchi R & Entling MH (2016) Effects of local and landscape factors on spiders and olive fruit flies. Agriculture, Ecosystems and Environment 222: 138-147.

Rader R, Bartomeus I, Garibaldi LA, Garratt MPD, Howlett B , Cunningham SA, Mayfield MM, Andersson GKS, Blanche R, Bommarco R, Brittain C, Carvalheiro LG, Chacoff NP, Entling MH, … and Woyciechowski M (2016) Non-bee insects are important contributors to global crop pollination. PNAS 113: 146-151.

Schirmel J, Bundschuh M, Entling MH, Kowarik I & Buchholz S (2016) Invasive plant effects on resident animals across ecosystems, taxa, and feeding types. A meta-analysis. Global Change Biology 22: 594-603.

Schirmel J, Thiele J, Entling MH & Buchholz S (2016) Trait composition and functional diversity of spiders and carabids in linear landscape elements. Agriculture, Ecosystems and Environment 235: 318-328.

Tschumi M, Albrecht M, Bärtschi C, Collatz J, Entling MH & Jacot K (2016) Perennial, species-rich wildflower strips enhance pest control and crop yield. Agriculture, Ecosystems and Environment 220: 97-103.

Tschumi M, Albrecht M, Collatz J, Dubsky V, Entling MH, Najar-Rodriguez AJ & Jacot K (2016) Tailored flower strips promote natural enemy biodiversity and enhance pest control in adjacent potato crops. Journal of Applied Ecology 53: 1169-1176.

Wagner J, Entling MH & Schirmel J (2016) Heuschrecken in biologisch und konventionell genutzten Weinbergen in der Pfalz. Articulata 31: 13-22.


Anjum-Zubair M, Entling MH, Bruckner A, Drapela T & Frank T (2015) Differentiation of carabid beetle assemblages between semi-natural habitats and adjoining winter wheat. Agricultural and Forest Entomology 17: 355-365.

Bucher R, Heinrich H & Entling MH (2015) Plant choice, herbivory and weight gain of wood crickets under the risk of predation. Entomologia Experimentalis et Applicata 155: 148-153.

Bucher R, Menzel F & Entling MH (2015) Impact of spider predation risk on arthropod communities and herbivory in the field. Oecologia 178: 571-577.

Buse J, Slachta M, Sladecek F, Carpaneto G & Entling MH (2015) Relative importance of pasture size and grazing continuity for the long-term conservation of European dung beetles. Biological Conservation 187: 112-119.

Buse J, Fassbender S, Entling MH & Pavlicek T (2015) Microclimatic divergence in a Mediterranean canyon affects richness, composition, and body size in saproxylic beetle assemblages. PLOS ONE 10: e0129323.

Hermann JD, Opatovsky I, Lubin Y, Pluess T, Gavish-Regev E & Entling MH (2015) Effects of non-native Eucalyptus plantations on epigeal spider communities in the northern Negev desert, Israel. Journal of Arachnology 43: 101-106.

Krell B, Röder N, Link M, Gergs R, Entling MH & Schäfer RB (2015) Aquatic prey subsidies to riparian spiders in a stream with different land use types. Limnologica 51: 1-7.

Pfister SC, Schäfer RB, Schirmel J & Entling MH (2015) Effects of hedgerows and riparian margins on aerial web-building spiders in cereal fields. Journal of Arachnology 43: 400-405.

Schulz R, Bundschuh M, Gergs R, Brühl CA, Diehl D, Entling MH, Fahse L, Frör O, Jungkunst HF, Lorke A, Schäfer RB, Schaumann GE & Schwenk K (2015) Review on environmental alterations propagating from aquatic to terrestrial ecosystems. Science of the Total Environment 538: 246-261.

Tschumi M, Albrecht M, Entling MH & Jacot K (2015) High effectiveness of tailored flower strips in reducing pests and crop plant damage. Proceedings of the Royal Society of London, Series B: Biological Sciences 282: 20151369.

Uhl P, Bucher R, Schäfer RB, Entling MH (2015) Sublethal effects of Imidacloprid in a tritrophic system. Chemosphere 132: 152-158.


Binz H, Bucher R, Entling MH & Menzel F (2014) Knowing the risk: crickets distinguish between spider predators of different size and commonness. Ethology 120: 99-110.

Bucher R, Binz H, Menzel F & Entling MH (2014) Spider cues stimulate feeding, weight gain and survival of crickets. Ecological Entomology 39: 667-673.

Bucher R, Binz H, Menzel F & Entling MH (2014) Effects of spider chemotactile cues on arthropod behaviour. Journal of Insect Behavior 27: 567-580.

Coudrain V, Schüepp C, Herzog F, Albrecht M & Entling MH (2014) Habitat amount modulates the effect of patch isolation on host-parasitoid interactions. Frontiers in Environmental Science 2: 27.

Hudson L, Newbold T, Contu S, Hill SLL, Lysenko I, ..., Entling MH, ..., Ewers RM, Mace GM, Purves DW, Scharlemann JPW & Purvis A (2014) The PREDICTS database: a global database of how local terrestrial biodiversity responds to human Impacts. Ecology and Evolution 4: 4701-4735.

Mestre L, Bucher R & Entling MH (2014) Trait-mediated effects between predators: ant chemical cues induce spider dispersal. Journal of Zoology 293: 119-125.

Schirmel J, Alt M, Rudolph I & Entling MH (2014). Effects of traditional flood irrigation on invertebrates in lowland meadows. PLOS ONE 9: e110854.

Schüepp C, Herzog F & Entling MH (2014) Disentangling multiple drivers of pollnation in a landscape-scale experiment. Proceedings of the Royal Society of London, Series B: Biological Sciences 281: 20132667.

Schüepp C, Uzman D, Herzog F & Entling MH (2014) Habitat isolation affects plant-herbivore-enemy interactions on cherry trees. Biological Control 71: 56-64.


Birkhofer K, Entling MH & Lubin Y (2013) Agroecology: Trait composition, spatial relationships, trophic interactions. Spider Research in the 21st Century (ed. D Penney), pp. 200-228. Siri Scientific Press, Manchester.

Buse J, Heinrich C, Weiter S & Entling MH (2013) Die Dungkäfer einer neu eingerichteten Waldweide bei St. Martin in der Pfalz. Mainzer Naturwissenschaftliches Archiv 50: 361-369.

Coudrain V, Herzog F & Entling MH (2013) Effects of habitat fragmentation on abundance, larval food and parasitism of a spider-hunting wasp. PLOS ONE 8: e59286.


Schüepp C, Rittiner S & Entling MH (2012) High bee and was diversity in a heterogeneous tropical farming system compared to protected forest. PLOS ONE 7: e52109.

Entling MH, Schweiger O, Bacher S, Espadaler X, Hickler T, Kumschick S, Woodcock BA & Nentwig W (2012) Species richness-environment relationships of European arthropods at two spatial grains: habitats and countries. PLOS ONE 7: e45875.

Entling MH (2012) Book Review: Invasive Alien Arthropod Predators and Parasitoids: An Ecological Approach (eds. H Roy, P De Clercq, L-J Lawson Handley, JJ Sloggett, RL Poland & E Wajnberg) Basic and Applied Ecology 13: 735-736.

Hermann JD, Kormann U, Schüepp C, Stocker Y, Herzog F & Entling MH (2012) Effects of habitat isolation and predation pressure on an arboreal food-web. Community Ecology 13: 82-87.


Bucher R & Entling MH (2011) Contrasting effects of habitat fragmentation, population density, and prey availability on body condition of two orb-weaving spiders. Ecological Entomology 36: 680-685.

Entling MH, Stämpfli K & Ovaskainen O (2011) Increased propensity for aerial dispersal in disturbed habitats due to intraspecific variation and species turnover. Oikos 120: 1099-1109.

Kumschick S, Fronzek S, Entling MH & Nentwig W (2011) Rapid spread of the wasp spider Argiope bruennichi across Europe: a consequence of climate change? Climatic Change 109: 319-329.

Sanders D & Entling MH (2011) Large variation of suction sampling efficiency depending on arthropod groups, species traits and habitat properties. Entomologia Experimentalis et Applicata 138: 234-243.

Schüepp C, Herrmann JD, Herzog F & Schmidt-Entling MH (2011) Differential effects of habitat isolation and landscape composition on wasps, bees, and their enemies. Oecologia 165: 713-721.

Stutz S & Entling MH (2011) Effects of the landscape context on aphid-ant-predator interactions on cherry trees. Biological Control 57: 37-43.

Hermann JD, Bailey D, Hofer G, Herzog F & Schmidt-Entling MH (2010) Spiders associated with the meadow and tree canopies of orchards respond differently to habitat fragmentation. Landscape Ecology 25: 1375-1384.

Bailey D, Schmidt-Entling MH, Eberhart P, Herrmann JD, Hofer G, Kormann U & Herzog F (2010) Effects of habitat amount and isolation on biodiversity in fragmented traditional orchards. Journal of Applied Ecology 47: 1003-1013.

Mühlner S, Kormann U, Schmidt-Entling MH, Herzog F & Bailey D (2010) Structural versus functional habitat connectivity measures to explain bird diversity in fragmented orchards. Journal of Landscape Ecology 3: 52-63.

Anjum-Zubair M, Schmidt-Entling MH, Querner P & Frank T (2010) Influence of within-field position and adjoining habitat type on carabid beetle assemblages in winter wheat. Agricultural and Forest Entomology 12: 301-306.

Bruggisser O, Schmidt-Entling MH & Bacher S (2010) Effects of vineyard management on biodiversity at three trophic levels. Biological Conservation 143: 1521-1528.

Entling W, Schmidt-Entling MH, Bacher S, Brandl R & Nentwig W (2010) Body size - climate relationships of European spiders. Journal of Biogeography 37: 233-237.

Pluess T, Opatovski I, Gavish-Regev E, Lubin Y & Schmidt-Entling MH (2010) Non-crop habitats in the landscape enhance spider diversity in wheat fields of a desert agroecosystem. Agriculture, Ecosystems and Environment 137: 68-74.

Opatovski I, Plüss T, Schmidt-Entling MH, Gavish-Regev E & Lubin Y. (2010) Are spider assemblages in fragmented, semi-desert habitat affected by increasing cover of agricultural crops? Agriculture, Ecosystems and Environment 135: 233-237.


Kumschick S, Schmidt-Entling MH, Bacher S, Hickler T, Entling W & Nentwig W (2009) Water limitation prevails over energy in European diversity gradients of sheetweb spiders (Araneae: Linyphiidae). Basic and Applied Ecology 10: 754-762.

Kumschick S, Schmidt-Entling MH, Bacher S, Hickler T, Espadaler X & Nentwig W (2009) Determinants of local ant (Hymenoptera: Formicidae) species richness and activity density across Europe. Ecological Entomology 34: 748-754.

Schmidt-Entling MH & Siegenthaler E (2009) Herbivore release through cascading risk effects. Biology Letters 5: 773-776.

Farwig N, Bailey D, Bochud E, Herrmann JD, Kindler E, Reusser N, Schuepp C & Schmidt-Entling MH (2009) Isolation from forest reduces pollination, seed predation and insect scavenging in Swiss farmland. Landscape Ecology 24: 919-927.

Schmidt-Entling MH & Döbeli J (2009) Sown wildflower areas to enhance spiders in arable fields. Agriculture, Ecosystems and Environment 133: 19-22.

Eichenberger B, Siegenthaler E & Schmidt-Entling MH (2009) Body size determines the outcome of competition for webs among alien and native sheetweb spiders (Araneae: Linyphiidae). Ecological Entomology 34: 363-368.

Muff P, Kropf C, Frick H, Nentwig W & Schmidt-Entling MH (2009) Coexistence of divergent communities at natural boundaries: spider (Arachnida: Araneae) diversity across an alpine timberline. Insect Conservation and Diversity 2: 36-44.


Jud P & Schmidt-Entling MH (2008) Fluid type, dilution, and bitter agent influence spider preservation in pitfall traps. Entomologia Experimentalis et Applicata 129: 356-359.

Schmidt MH, Rocker S, Hanafi J & Gigon A (2008) Rotational fallows as overwintering habitat for grassland arthropods: the case of spiders in fen meadows. Biodiversity and Conservation 17: 3003-3012.

Pluess T, Opatovsky I, Gavish-Regev E, Lubin Y & Schmidt MH (2008) Spiders in wheat fields and semi-desert in the Negev (Israel). Journal of Arachnology 36: 368-373.

Schmidt MH, Thies C, Nentwig W & Tscharntke T (2008) Contrasting responses of arable spiders to the landscape matrix at different spatial scales. Journal of Biogeography 35: 157-166.


Entling W, Schmidt MH, Bacher S, Brandl R & Nentwig W (2007) Niche properties of Central European spiders: shading, moisture, and the evolution of the habitat niche. Global Ecology and Biogeography 18: 440-448.

Muff P, Schmidt MH, Frick H, Nentwig W & Kropf C (2007) Spider (Arachnida: Araneae) distribution across the timberline in the Swiss Central Alps (Alp Flix, Grisons) and three morphologically remarkable species. Arachnologische Mitteilungen 34: 16-24.

Schmidt MH & Hänggi A (2007) Zelotes mundus (Araneae: Gnaphosidae) in the Camargue: a continental species meets the Mediterranean coast. Bulletin of the British Arachnological Society 14: 27-29.


Schmidt MH, Clough Y, Schulz W, Westphalen A & Tscharntke T (2006) Capture efficiency and preservation attributes of different fluids in pitfall traps. Journal of Arachnology 34: 159-162.

Flückiger R & Schmidt MH (2006): Contribution of sown wildflower areas to cereal aphid control: from local to landscape scale. IOBC/WPRS Bulletin 29: 41-44.


Schmidt MH, Roschewitz I, Thies C & Tscharntke T (2005) Differential effects of landscape and management on diversity and density of ground-dwelling farmland spiders. Journal of Applied Ecology 42: 281-287.

Schmidt MH & Tscharntke T (2005) Landscape context of sheetweb spider (Araneae: Linyphiidae) abundance in cereal fields. Journal of Biogeography 32: 467-473.

Schmidt MH & Tscharntke T (2005) The role of perennial habitats for Central European farmland spiders. Agriculture, Ecosystems and Environment 105: 235-242.

Schmidt MH, Lefebvre G, Poulin B & Tscharntke T (2005) Reed cutting affects arthropod communities, potentially reducing food for passerine birds. Biological Conservation 121: 157-166.


Schmidt MH, Thewes U, Thies C & Tscharntke T (2004) Aphid suppression by natural enemies in mulched cereals. Entomologia Experimentalis et Applicata 113: 87-93.

Schmidt MH, Thies C & Tscharntke T (2004) The landscape context of arthropod biological control. Ecological Engineering for Pest Management: Advances in Habitat Manipulation for Arthropods (eds. GM Gurr, SD Wratten & MA Altieri), pp. 55-63. CSIRO, Collingwood VIC.


Schmidt MH, Lauer A, Purtauf T, Thies C, Schaefer M & Tscharntke T (2003) Relative importance of predators and parasitoids for cereal aphid control. Proceedings of the Royal Society of London, Series B: Biological Sciences 270: 1905-1909.


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AGROMIX - AGROforestry and MIXed farming systems - Participatory research to drive the transition to a resilient and efficient land use in Europe

FRAGMENT III: Effects of spatio-temporal resource availability on pollinators and pest-natural enemies in fragmented agricultural landscapes

GETREAL: Incorporating spatial and temporal variability in community sensitivity into chemical risk assessment

MonViA: Hymenoptera in viticulture

Permaculture: Biodiversity, Soil Quality, Carbon Storage and Resource Efficiency of Permaculture Farms in Central Europe

PiWiDiv: Effects of fungicide reduction on biodiversity in vineyards

SECBIVIT: Scenarios for providing multiple ecosystem services and biodiversity in viticultural landscapes

SystemLink: Aqutic subsidies in terrestrial food webs

SystemLink: The response of riparian food webs to chemical pollutants in streams


Ökologie, Biodiversität und Evolution

Conservation Biology

Angewandte Ökologie

Physiologie und Ökologie der Tiere

Indicator Organisms: Spiders



iES Landau - Institut für Umweltwissenschaften